:root{--ink:#142e3a;--muted:#597078;--navy:#09232e;--teal:#0e716d;--paper:#fbfbf7;--line:#d8e0dc;--sea:#dbe9e4;--serif:Georgia,'Times New Roman',serif;--sans:Arial,Helvetica,sans-serif}*{box-sizing:border-box}html{scroll-behavior:smooth;scroll-padding-top:24px}body{margin:0;background:var(--paper);color:var(--ink);font:16px/1.6 var(--sans)}h1,h2,h3,p,figure{margin:0}a{color:inherit;text-decoration:none}a:hover{text-decoration:underline;text-underline-offset:4px}button,input,select,textarea{font:inherit}button,a,input,select,textarea{touch-action:manipulation}button{cursor:pointer}img{display:block;max-width:100%;height:auto}a:focus-visible,button:focus-visible,input:focus-visible,select:focus-visible,textarea:focus-visible{outline:3px solid #d99736;outline-offset:4px}.sr-only{position:absolute;width:1px;height:1px;padding:0;overflow:hidden;clip:rect(0,0,0,0);white-space:nowrap}.skip{position:absolute;top:-80px;left:20px;background:white;padding:10px;z-index:20}.skip:focus{top:10px}.section,.mast-inner,.nav-inner,.edition-strip,.footer-top,.footer-bottom{width:min(1180px,calc(100% - 80px));margin:auto}.masthead{background:var(--paper)}.mast-inner{height:111px;display:flex;align-items:center;gap:35px}.brand{display:flex;align-items:center;gap:13px;font-size:25px;font-weight:700;line-height:1.1;letter-spacing:1.2px}.brand:hover{text-decoration:none}.brand-sub{display:block;font-size:11px;letter-spacing:5.3px;margin-top:9px}.tagline{border-left:1px solid var(--line);padding-left:30px;font:italic 15px/1.5 var(--serif);color:var(--muted)}.header-search{display:flex;margin-left:auto;border-bottom:1px solid #9faeac;max-width:245px}.header-search input{background:none;border:0;padding:9px 0;width:192px;font-size:13px;color:var(--ink)}.header-search button{border:0;background:none;font-size:23px;color:var(--teal);padding:0 5px}.masthead nav{border-top:1px solid var(--line);border-bottom:1px solid var(--line)}.nav-inner{display:flex;gap:39px;align-items:center;min-height:49px;font-size:13px;font-weight:600}.nav-inner a{padding:13px 0;position:relative}.nav-inner a[aria-current]{color:var(--teal)}.nav-inner a[aria-current]:after{content:'';height:3px;position:absolute;bottom:0;background:var(--teal);left:0;right:0}.nav-end{margin-left:auto;color:var(--muted);font-weight:400}.hero{height:490px;position:relative;overflow:hidden;background:#020a11;color:white}.hero-image{position:absolute;width:100%;height:100%;object-fit:cover;object-position:center 53%}.hero:after{content:'';position:absolute;inset:0;background:linear-gradient(90deg,rgba(2,14,23,.88),rgba(2,14,23,.53) 37%,transparent 70%);pointer-events:none}.hero-inner{width:min(1180px,calc(100% - 80px));position:relative;margin:0 auto;padding-top:54px;z-index:1}.eyebrow{font:600 10px/1.5 var(--sans);letter-spacing:1.8px;text-transform:uppercase;color:var(--teal)}.hero .eyebrow{color:#a3d3c9;font-size:10px}.hero h1{font:normal 59px/1.08 var(--serif);letter-spacing:-1.4px;margin:20px 0}.hero-deck{font:17px/1.6 var(--serif);color:#d5e1e3}.button{display:inline-flex;align-items:center;justify-content:space-between;gap:30px;padding:12px 20px;border:1px solid transparent;font:600 12px/1.4 var(--sans);border-radius:0;min-height:43px}.button:hover{text-decoration:none;filter:brightness(.95)}.button.light{background:#e4eee9;color:#133d42;margin-top:25px}.button.dark{background:var(--teal);color:white}.hero-feature{display:block;font-size:11px;margin-top:28px;color:#d6e5e6}.hero-feature span{font-size:9px;letter-spacing:1.5px;color:#90b9b8;margin-right:12px}.image-credit{position:absolute;z-index:2;bottom:19px;right:30px;color:#b3c0c9;font-size:9px}.edition-strip{display:flex;align-items:center;gap:24px;min-height:56px;font-size:11px;border-bottom:1px solid var(--line)}.edition-strip>span{font-size:9px;font-weight:700;letter-spacing:1.5px;color:var(--teal)}.edition-strip p{font:italic 13px var(--serif);color:var(--muted)}.edition-strip a{margin-left:auto;color:var(--teal)}.organisms{padding-top:62px;padding-bottom:54px}.section-heading{display:flex;align-items:end;justify-content:space-between;gap:25px;margin-bottom:19px}.section-heading h2,.depth-copy h2,.field-band h2{font:normal 34px/1.2 var(--serif);letter-spacing:-.5px;margin-top:8px}.section-heading>a{font-size:12px;color:var(--teal);padding-bottom:3px;white-space:nowrap}.section-intro{font-size:14px;color:var(--muted);max-width:660px;margin-bottom:30px}.organism-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:36px 28px}.specimen{position:relative;background:#0b2430;overflow:hidden}.specimen img{width:100%;aspect-ratio:1.62;object-fit:contain}.specimen-number{position:absolute;top:13px;left:16px;color:#94b8bb;font:10px var(--sans)}.organism:hover .specimen img{transform:scale(1.025)}.specimen img{transition:transform .25s}.organism .taxon{display:block;font-size:9px;text-transform:uppercase;letter-spacing:1.7px;color:var(--teal);margin-top:18px}.organism h3,.organism h2{font:normal 25px/1.25 var(--serif);margin:5px 0 8px}.organism p{color:var(--muted);font-size:13px;line-height:1.65}.text-link{display:block;font-size:11px;color:var(--teal);font-weight:600;margin-top:14px}.organism:hover,.story a:hover{text-decoration:none}.organism:hover h3,.story a:hover h3,.organism a:hover h2{text-decoration:underline;text-underline-offset:4px}.figure-note{font-size:10px;line-height:1.6;color:var(--muted);margin-top:23px}.journal-section{background:#eef1eb;border-block:1px solid var(--line)}.journal-section>.section{padding-block:49px 52px}.story-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:28px}.story{min-width:0}.story img{width:100%;background:#0c2834;aspect-ratio:1.85;object-fit:contain}.story a>div,.collection-copy{padding-top:20px}.story h3,.story h2{font:normal 25px/1.22 var(--serif);margin:9px 0 12px}.story p:not(.eyebrow){font-size:13px;line-height:1.75;color:var(--muted)}.story .text-link{margin-top:18px}.depth-section{display:grid;grid-template-columns:1fr 1fr;gap:85px;padding-block:62px}.depth-copy>p:not(.eyebrow):not(.figure-note){max-width:420px;font-size:14px;color:var(--muted);margin-top:18px}.depth-buttons{display:flex;gap:7px;margin-top:23px;flex-wrap:wrap}.depth-buttons button{background:transparent;border:1px solid #bbc9c5;color:var(--ink);padding:8px 12px;font-size:12px}.depth-buttons button[aria-pressed=true]{background:var(--teal);border-color:var(--teal);color:white}.depth-description{min-height:125px;margin-top:22px;max-width:420px}.depth-description h3{font:22px var(--serif);margin-bottom:7px}.depth-description p{font-size:13px;color:var(--muted)}.depth-copy>a{font-size:12px;color:var(--teal)}.depth-copy h2{font-size:37px}.depth-visual{background:linear-gradient(#438c98,#123d54 31%,#091d32 60%,#040c1a);position:relative;min-height:430px;overflow:hidden}.depth-surface{height:8px;background:repeating-linear-gradient(120deg,#b0d7d0 0,#4a99a1 20px,#b0d7d0 45px)}.zone-label{position:absolute;left:22px;color:#bad3d5;font:9px var(--sans);letter-spacing:2px}.label-1{top:30px}.label-2{top:150px}.label-3{top:286px}.depth-marker{position:absolute;left:0;right:0;top:50px;transition:top .6s;border-top:1px dashed #91c8c3;z-index:2}.depth-marker span{position:absolute;right:18px;top:6px;font-size:12px;color:#e5efeb;background:#143a48;padding:1px 7px}.depth-marker[data-level="200"]{top:155px}.depth-marker[data-level="1000"]{top:280px}.depth-marker[data-level="4000"]{top:375px}.depth-animal{position:absolute;top:90px;left:100px;right:20px;opacity:.7}.depth-animal img{width:100%}.depth-bottom{position:absolute;bottom:17px;left:22px;color:#8faab2;font-size:9px}.field-band{background:#e2eae2;border-top:1px solid #cfdbd4}.field-band>.section{display:grid;grid-template-columns:1.3fr 1fr;gap:100px;padding-block:43px 46px}.field-band h2{font-size:30px;margin-top:10px}.field-band p:not(.eyebrow){font-size:13px;max-width:470px;margin-top:17px;color:var(--muted)}.field-band .button{margin-top:24px}.field-band ul{list-style:none;padding:0;margin:17px 0 0}.field-band li{border-bottom:1px solid #bdcec3}.field-band li a{padding:16px 0;display:flex;justify-content:space-between;gap:20px;font:17px var(--serif)}.field-band li b{font:normal 20px var(--sans);color:var(--teal)}footer{background:var(--navy);color:#d7e2e0;margin-top:0}.footer-top{display:flex;gap:60px;padding-block:46px}.footer-brand{font-size:20px;font-weight:700;letter-spacing:1px;line-height:1.4}.footer-brand span{display:block;font-size:9px;letter-spacing:4px;margin-top:5px}.footer-top>p{font:italic 16px/1.6 var(--serif);color:#adbfbd}.footer-top>div{display:grid;grid-template-columns:1fr 1fr;gap:8px 35px;font-size:11px;margin-left:auto}.footer-bottom{border-top:1px solid #25414b;padding-block:20px;display:flex;justify-content:space-between;gap:20px;font-size:9px;color:#a4b9bc}.page-head{padding-block:50px 36px;border-bottom:1px solid var(--line)}.page-head h1{font:normal 50px/1.12 var(--serif);letter-spacing:-1px;margin:15px 0}.page-head>p:last-child{max-width:720px;font:19px/1.55 var(--serif);color:var(--muted)}.collection{padding-block:38px 60px}.collection .organism-grid{row-gap:45px}.collection .collection-copy{padding-top:18px}.collection h2{font-size:26px}.article-head{padding-block:35px;max-width:1180px}.breadcrumbs{font-size:11px;color:var(--muted);margin-bottom:30px}.breadcrumbs a{text-decoration:underline}.article-head h1{font:normal 49px/1.12 var(--serif);letter-spacing:-1px;max-width:930px;margin:13px 0 20px}.article-deck{font:20px/1.55 var(--serif);max-width:780px;color:var(--muted)}.byline{font-size:10px;color:var(--muted);margin-top:24px}.article-figure{width:min(1000px,calc(100% - 80px));margin:0 auto 0;background:#0b2430}.article-figure img{width:100%;height:390px;object-fit:contain}.article-figure figcaption{background:var(--paper);padding:12px 0;font-size:11px;color:var(--muted)}.article-layout{display:grid;grid-template-columns:220px minmax(0,700px);gap:65px;padding-block:37px 60px;justify-content:center}.article-nav{font-size:12px}.article-nav ol{padding-left:18px;line-height:1.6}.article-nav li{margin-bottom:12px}.taxonomy{border-top:1px solid var(--line);margin-top:30px;padding-top:18px}.taxonomy span{font-size:9px;letter-spacing:1.7px;color:var(--muted)}.taxonomy strong{display:block;font:22px var(--serif);margin:8px 0}.taxonomy p{color:var(--muted);font-size:11px}.article-body{font:18px/1.8 var(--serif);min-width:0}.article-body h2,.prose-page h2{font:normal 28px/1.25 var(--serif);margin:31px 0 16px}.article-body section:first-of-type h2{margin-top:0}.article-body p{margin-bottom:19px}.article-body a,.prose-page a{color:var(--teal);text-decoration:underline;text-underline-offset:3px;overflow-wrap:anywhere}.article-body .citation{font:11px var(--sans);vertical-align:super;text-decoration:none}.article-body .restoration-note{font:11px/1.7 var(--sans);border-left:2px solid #83aaa0;padding-left:14px;color:var(--muted);margin-bottom:27px}.references{border-top:1px solid var(--line);margin-top:35px;padding-top:4px}.references h2{font-size:23px}.references ol{font:13px/1.7 var(--sans);padding-left:20px}.references li{margin-bottom:10px;scroll-margin-top:25px}.references li:target{background:#e0e9df}.correction-link{font:11px/1.7 var(--sans);color:var(--muted);border-top:1px solid var(--line);margin-top:28px;padding-top:20px}.related{padding-bottom:50px}.related-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:25px;margin-top:15px}.related-grid a{font:20px/1.4 var(--serif);border-top:1px solid var(--line);padding-top:16px}.related-grid span{color:var(--teal)}.library-layout{display:grid;grid-template-columns:240px minmax(0,1fr);gap:60px;padding-block:37px 60px}.library-search{display:grid;gap:10px}.library-search label{font-size:12px;font-weight:600;margin-top:6px}.library-search input,.library-search select,.contact-form input,.contact-form textarea{width:100%;background:#fff;border:1px solid #b9c9c3;padding:10px;color:var(--ink);border-radius:0;font-size:14px}.library-search .button{margin-top:8px}.library-search>a{font-size:12px;color:var(--teal)}.result-count{font-size:11px;color:var(--muted);padding-bottom:13px;border-bottom:1px solid var(--line)}.reading-list{list-style:none;padding:0;margin:0}.reading-list li{padding:24px 0;border-bottom:1px solid var(--line)}.reading-list h2{font:normal 25px/1.3 var(--serif);margin:7px 0 10px}.reading-list h2 span{font:18px var(--sans);color:var(--teal)}.reading-list li>p:last-child{font-size:13px;line-height:1.75;color:var(--muted)}.pagination{display:flex;align-items:center;justify-content:center;gap:25px;margin-top:30px;font-size:12px}.pagination a{color:var(--teal)}.prose-page{max-width:760px;padding-block:35px 65px;font:18px/1.85 var(--serif)}.prose-page p{margin-bottom:20px}.prose-page .lead{font-size:23px;line-height:1.6;color:#34555b}.prose-page h2{margin-top:30px}.source-list{padding-left:24px;font:15px/1.7 var(--sans)}.source-list li{margin-block:18px}.source-list span{display:block;font-size:11px;color:var(--muted)}.contact-form{display:grid;gap:12px;font:14px/1.7 var(--sans)}.contact-form label{font-weight:600;margin-top:10px}.contact-form p{font-size:12px;color:var(--muted)}.contact-form textarea{resize:vertical}.contact-form .button{justify-self:start}.success{border-left:3px solid var(--teal);padding-left:22px}.form-error{color:#9c362b}
@media(min-width:1600px){.hero{height:540px}.hero-inner{padding-top:70px}.hero-image{object-position:center 58%}.hero h1{font-size:67px}}
@media(max-width:1000px){.section,.mast-inner,.nav-inner,.edition-strip,.footer-top,.footer-bottom,.hero-inner{width:calc(100% - 48px)}.nav-inner{gap:25px}.tagline{font-size:13px;padding-left:20px}.mast-inner{gap:24px}.hero h1{font-size:54px}.hero{height:475px}.organism-grid{gap:28px 21px}.depth-section{gap:40px}.field-band>.section{gap:50px}.article-layout{gap:35px;grid-template-columns:180px minmax(0,1fr)}.library-layout{grid-template-columns:200px minmax(0,1fr);gap:35px}.footer-top{gap:35px}.footer-top>div{column-gap:25px}}
@media(max-width:700px){.section,.mast-inner,.nav-inner,.edition-strip,.footer-top,.footer-bottom,.hero-inner{width:calc(100% - 36px)}.mast-inner{height:88px;gap:16px}.brand{font-size:21px;gap:10px}.brand img{width:35px}.brand-sub{font-size:9px;letter-spacing:4px;margin-top:7px}.tagline{display:none}.header-search{width:38px;border:0}.header-search input{position:absolute;width:1px;height:1px;clip:rect(0,0,0,0)}.header-search button{font-size:27px;width:36px}.nav-inner{gap:0;justify-content:space-between;font-size:11px;min-height:46px}.nav-end{display:none}.nav-inner a{padding:13px 0}.hero{height:505px}.hero-image{object-fit:cover;object-position:62% 10%;opacity:.8}.hero:after{background:linear-gradient(90deg,rgba(2,14,23,.86),rgba(2,14,23,.24)),linear-gradient(0deg,rgba(2,14,23,.65),transparent 65%)}.hero-inner{padding-top:42px}.hero h1{font-size:48px;line-height:1.12;margin-top:25px}.hero .eyebrow{font-size:8px;letter-spacing:1.5px}.hero-deck{font-size:16px}.hero .button{margin-top:24px}.hero-feature{margin-top:34px;font-size:10px;max-width:280px}.hero-feature span{display:block;margin-bottom:5px}.image-credit{font-size:8px;right:18px;bottom:14px}.edition-strip{padding-block:15px;display:grid;grid-template-columns:1fr;gap:3px}.edition-strip a{display:none}.edition-strip p{font-size:12px}.organisms{padding-block:38px}.section-heading{align-items:start;gap:15px}.section-heading h2{font-size:29px}.section-heading>a{font-size:10px;margin-top:24px;max-width:95px;white-space:normal;text-align:right}.section-intro{font-size:13px;margin-bottom:22px}.organism-grid{grid-template-columns:repeat(2,minmax(0,1fr));gap:27px 17px}.organism h3,.organism h2{font-size:22px}.organism p{font-size:12px}.organism .taxon{font-size:8px;letter-spacing:1.1px;margin-top:12px}.specimen img{aspect-ratio:1.25}.specimen-number{top:9px;left:10px;font-size:8px}.text-link{font-size:10px}.figure-note{font-size:9px}.journal-section>.section{padding-block:33px}.story-grid{grid-template-columns:1fr;gap:28px}.story a{display:grid;grid-template-columns:125px 1fr;gap:17px}.story a>img{height:125px;aspect-ratio:1;object-fit:contain}.story a>div{padding-top:0}.story h3,.story h2{font-size:23px;margin-top:6px}.story .eyebrow{font-size:8px}.story p:not(.eyebrow){font-size:12px}.story .text-link{margin-top:10px}.depth-section{grid-template-columns:1fr;gap:30px;padding-block:38px}.depth-copy h2{font-size:33px}.depth-description{min-height:0;margin-bottom:20px}.depth-visual{min-height:410px;max-width:500px;width:100%}.depth-animal{left:80px}.field-band>.section{grid-template-columns:1fr;gap:15px;padding-block:33px}.field-band h2{font-size:29px}.field-band li a{font-size:17px}.footer-top{flex-wrap:wrap;gap:25px;padding-block:32px}.footer-top>p{font-size:14px;flex:1;min-width:140px}.footer-top>div{width:100%;margin:0;grid-template-columns:1fr 1fr;font-size:11px}.footer-bottom{display:block;font-size:9px;padding-block:17px}.footer-bottom>a{display:block;margin-top:7px}.page-head{padding-block:34px 26px}.page-head h1{font-size:39px}.page-head>p:last-child{font-size:17px}.collection{padding-block:28px 42px}.collection .story a{display:block}.collection .story a>div{padding-top:15px}.collection .story a>img,.collection .story .specimen img{width:100%;height:auto;aspect-ratio:1.8}.collection .organism h2{font-size:22px}.article-head{padding-block:25px}.breadcrumbs{font-size:10px;margin-bottom:23px}.article-head h1{font-size:37px;line-height:1.17;letter-spacing:-.5px}.article-deck{font-size:18px}.article-figure{width:calc(100% - 36px)}.article-figure img{height:240px}.article-figure figcaption{font-size:10px}.article-layout{display:block;padding-top:23px}.article-nav{margin-bottom:28px;border-block:1px solid var(--line);padding-block:17px}.article-nav ol{display:flex;flex-wrap:wrap;gap:8px 23px;padding-left:18px;margin-bottom:0}.article-nav li{margin:0;width:calc(50% - 23px);font-size:10px}.taxonomy{display:none}.article-body{font-size:17px;line-height:1.8}.article-body h2{font-size:26px}.related-grid{grid-template-columns:1fr;gap:20px}.related-grid a{font-size:22px}.library-layout{display:block;padding-block:25px 40px}.library-layout>aside{margin-bottom:30px}.library-search{grid-template-columns:1fr 1fr;gap:9px 13px}.library-search label{grid-row:1;font-size:11px}.library-search input,.library-search select{grid-row:2;min-width:0;font-size:12px}.library-search .button{grid-column:1/3;justify-content:center;margin-top:5px}.library-search>a{grid-column:1/3}.reading-list h2{font-size:23px}.prose-page{padding-block:28px 45px;font-size:17px}.prose-page .lead{font-size:21px}.prose-page h2{font-size:26px}}
@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}*,*:before,*:after{transition:none!important}}
/* Source-led reading paths, photographic references and reproducible data. */
.topic-shortcut{font:13px/1.7 var(--sans);border-left:3px solid var(--teal);padding:8px 14px;margin:0 0 24px}.photo-evidence{margin-bottom:30px}.photo-evidence img{width:100%;height:auto;max-height:580px;object-fit:contain;background:#0b2430}.photo-evidence figcaption,.data-case figcaption{font:11px/1.7 var(--sans);color:var(--muted);margin-top:10px}.table-scroll{overflow-x:auto;margin:22px 0}.table-scroll table{width:100%;border-collapse:collapse;min-width:470px;font:13px/1.6 var(--sans)}.table-scroll th,.table-scroll td{text-align:left;vertical-align:top;padding:12px;border-bottom:1px solid var(--line)}.table-scroll thead{background:#e5ece6}.table-scroll caption{text-align:left;font-size:12px;margin-bottom:8px}.worksheet{border:1px solid var(--line);padding:20px;background:#edf1eb;margin:25px 0}.worksheet h2{margin-top:0!important;font-size:24px}.worksheet p,.worksheet a{font:14px/1.7 var(--sans)}.data-case{border-block:1px solid var(--line);padding:24px 0;margin:24px 0}.data-case details{font:14px/1.7 var(--sans);margin:20px 0}.data-case summary{cursor:pointer;color:var(--teal);font-weight:600}.data-case h3{font:23px/1.4 var(--serif);margin:25px 0 12px}.article-body>section+section{margin-top:30px}.data-case figure img{width:100%;height:auto}.byline a{text-decoration:underline}.editorial-link{font-size:11px;color:var(--muted)}

.original-work{display:grid;grid-template-columns:1fr 1.25fr;gap:50px;padding-block:42px;border-top:1px solid var(--line)}.original-work h2{font:32px/1.25 var(--serif);margin:10px 0 18px}.original-work p:not(.eyebrow){font-size:14px;color:var(--muted)}.original-work>a{align-self:center}.original-work img{width:100%}@media(max-width:700px){.original-work{grid-template-columns:1fr;gap:25px}.original-work h2{font-size:29px}}
